1. Graduate School of Animal and Veterinary Sciences and Agriculture, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan
2. Research Center for Global Agromedicine, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan
3. Graduate School of Agricultural Science, Kobe University, Kobe, Hyogo, Japan
4. Graduate School of Agriculture, Hokkaido University, Sapporo, Hokkaido, Japan
5. Department of Statistics and Department of Plant Pathology, University of Wisconsin-Madison, Madison, WI, USA