1. Department of Watershed Sciences and The Ecology Center, Utah State University, 5210 Old Main Hill, Logan, Utah 84322, USA
2. U.S. Geological Survey, Utah Cooperative Fish and Wildlife Research Unit, 5290 Old Main Hill, Logan, Utah 84322, USA; and Department of Watershed Science and The Ecology Center, Utah State University, 5210 Old Main Hill, Logan, Utah 84322, USA
3. Southeast Environmental Research Center and Department of Biological Sciences, Florida International University, 11200 Southwest 8th Street, Miami, Florida 33199, USA