1. Department of Biological Sciences, Michigan Technological University, 1400 Townsend Drive, Houghton, Michigan 49931, USA
2. Grand Rapids Public Museum, 272 Pearl Street Northwest, Grand Rapids, Michigan 49504, USA
3. Michigan Department of Natural Resources, Fisheries Division, Tribal Coordination Unit, Charlevoix Fisheries Research Station, 96 Grant Street, Charlevoix, Michigan 49720, USA