1. School of Aquatic and Fishery Sciences, University of Washington, Box 355020, Seattle, Washington 98195, USA
2. National Oceanic and Atmospheric Administration, National Marine Fisheries Service, Northwest Fisheries Science Center, Environmental and Fisheries Sciences Division, 2725 Montlake Boulevard East, Seattle, Washington 98112, USA
3. Biological Sciences Department, Center for Coastal Marine Sciences, California Polytechnic State University, San Luis Obispo, California 93407, USA