1. School of Freshwater Sciences, University of Wisconsin–Milwaukee, 600 East Greenfield Avenue, Milwaukee, Wisconsin 53204, USA
2. Michigan Department of Natural Resources, Marquette Fisheries Research Station, 484 Cherry Creek Road, Marquette, Michigan 49855, USA
3. School of Aquatic and Fishery Science, University of Washington, Box 355020, Seattle, Washington 98195-5020, USA
4. U.S. Fish and Wildlife Service, Ashland Fish and Wildlife Conservation Office, 2800 Lake Shore Drive East, Ashland, Wisconsin 54806, USA