1. Red Sea Research Center, Division of Biological and Environmental Science and Engineering, King Abdullah University of Science and Technology, Thuwal, 23955–6900, Saudi Arabia
2. ARC Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, QLD, 4811, Australia
3. Department of Biotechnologies and Bioscience, University of Milano – Bicocca, Milan, 20126, Italy
4. European Commission, Joint Research Centre, Directorate A – Strategy, Work Programme and Resources, Exploratory Research, Ispra, 21027, Italy