1. Faculty of Fisheries, Kagoshima University, 4-50-20 Shimoarata, Kagoshima 890-0056, Japan
2. Department of Biological and Environmental Sciences, Georgia College & State University, Campus Box 081, Milledgeville, GA 31061, USA
3. Ecological Genetics Laboratory, National Institute of Genetics, 1111 Yata, Mishima, Shizuoka 411-8540, Japan
4. Department of Genetics, The Graduate University for Advanced Studies, Mishima, Shizuoka, Japan, 1111 Yata, Mishima, Shizuoka 411-8540, Japan
5. Tateyama Marine Laboratory, Institute for Marine and Coastal Research, Ochanomizu University, 11 Kouyatsu, Tateyama, Chiba 294-0301, Japan