1. Graduate Degree Program, Division of EcoScience, Ewha Womans University, 52 Ewhayeodae-gil, Seodaemun-gu, Seoul, 03760, South Korea
2. Present Address: Nakdonggang National Institute of Biological Resources, Sangju-si, Gyeongsangbuk-do, 37242, South Korea
3. Division of EcoScience, Ewha Womans University, 52 Ewhayeodae-gil, Seodaemun-gu, Seoul, 03760, South Korea
4. Present Address: Graduate Degree Program, Department of Ecology and Evolutionary Biology, Yale University, P.O.Box 208106, New Haven, Connecticut, 06520-8106, USA
5. Inland Fisheries Research Institute, National Fisheries Research and Development Institute, 65 Gangbyeon-ro, Gapyeong-gun, Gyeonggi-do, 12453, South Korea