1. Department of Evolutionary Biology, Ecology and Environmental Sciences (Arthropods), Biodiversity Research Institute (IRBio), University of Barcelona, Avd. Diagonal, 643, Barcelona, 08028, Spain
2. Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History, Helsinki, 00014, Finland
3. Instituto das Florestas e Conservação da Natureza, Funchal, 9054-505, Portugal
4. Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History, P.O. Box 17, Helsinki, 00014, Finland
5. University of Helsinki, Helsinki, 00014, Finland