1. a Faculty of Science , University of the Ryukyus, Senbaru 1 , Nishihara , Okinawa , 903-0213 , Japan
2. b Center for Advanced Studies of Blanes , Spanish National Research Council (CEAB-CSIC) , Blanes , Girona , Spain
3. c Department of Animal Biology (Invertebrates) , University of Barcelona, 643 Diagonal Avenue , 08028 , Barcelona , Spain