1. Molecular Invertebrate Systematics and Ecology Laboratory, Graduate School of Engineering and Science, University of the Ryukyus, Senbaru 1, Nishihara, Okinawa, 903-0213, Japan
2. Centro de Ciências do Mar, Universidade do Algarve, Faro, P-8005-139, Portugal
3. Tropical Biosphere Research Center, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa, 903-0213, Japan
4. Departamento de Biología Animal, Edafología y Geología, Universidad de La Laguna (ULL), Avenida Astrofísico Francisco Sánchez, s/n, La Laguna, Tenerife, 38206, Spain
5. Marine Section, Environmental Management Division, Saint Helena Government