1. Muséum national d’Histoire naturelle, CNRS, SU, EPHE, UA, Institut de Systématique, Evolution et Biodiversité (ISYEB), 57 rue Cuvier, CP 50, Paris Cedex 05, 75231, France
2. Zoological Research Museum Alexander Koenig, Adenauerallee 160, Bonn, D-53113, Germany
3. Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE1410, Brunei Darussalam
4. Sabah Forestry Department, Forest Research Centre (Sepilok), P.O. Box 1407, Sandakan, 90715, Sabah