1. Natural History Museum and Institute, Chiba. Aoba-cho 955-2, Chuo-ku, Chiba, 260-8682, Japan
2. Department of Biology, Toho University, 2-2-1 Miyama, Funabashi, 274-8510, Chiba, Japan
3. Research Development and Innovation Division, Section of Biodiversity Fauna, Forest Department Sarawak, Kuching, 93250, Sarawak, Malaysia
4. Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu-cho, Sakyo-ku, Kyoto, 606-8501, Japan
5. Graduate School of Global Environmental Studies, Kyoto University, Yoshida Hon-machi, Sakyo-ku, Kyoto, 606-8501, Japan