1. Bioscience group, National Institute of Polar Research, 10-3 Midori-cho, Tachikawa, 190-8518, Tokyo, Japan
2. Graduate School of Bioresources, Mie University, 1577 Kurimamachiya-cho, Tsu, 514-8507, Mie, Japan
3. Graduate School of Science, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo, 113-0033, Japan
4. Faculty of Science, Hokkaido University, Kita 10 Nishi 8 Kitaku, Sapporo, 060-0810, Hokkaido, Japan