1. Department of Biological Sciences, Macquarie University, Sydney, Australia
2. Fowlers Gap Arid Zone Research Station, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, Australia
3. Centre for Ecology and Conservation, College of Life and Environmental Sciences, University of Exeter, Penryn, UK
4. School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, Australia