1. Research Institute for the Environment and Livelihoods, Charles Darwin University, Casuarina, NT, Australia
2. Biodiversity and Conservation Science, Department of Biodiversity, Conservation and Attractions, Bentley, WA, Australia
3. School of Science, Edith Cowan University, Joondalup, WA, Australia
4. Department of Ecology and Evolution, University of Chicago, Chicago, IL, USA