1. Department of Geology and Palaeontology, Muséum d’Histoire naturelle, CP 6434, 1211 Genève 6, Switzerland
2. Department of Biology, Faculty of Science, Mahasarakham University, Khamrieng, Kantharawichai District, Mahasarakham 44150, Thailand,
3. Palaeontological Research and Education Centre, Mahasarakham University, Khamrieng, Kantharawichai District, Mahasarakham 44150, Thailand
4. Sirindhorn Museum, Department of Mineral Resources, 200 Moo 11, Nonburi, Kalasin, 46140, Thailand,