1. Department of Zoology and Entomology, University of the Free State, Bloemfontein, 9300, South Africa
2. Evolutionary Studies Institute University of the Witwatersrand, Johannesburg, South Africa,
3. School of Geosciences, University of the Witwatersrand, Johannesburg, South Africa
4. National Museum, PO Box 266, Bloemfontein, 9300, South Africa, .