1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China,
2. Chinese Academy of Sciences Center for Excellence in Life and Paleoenvironment, Beijing 100044, China
3. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing 100049, China,
4. American Museum of Natural History, New York, New York 10024, U.S.A.,