1. Biodiversity Institute and Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, U.S.A.
2. Centre de Recherche en Paléontologie–Paris, UMR 7207 (CNRS, MNHN, UPMC), Sorbonne Université, Muséum National d’Histoire Naturelle, 8 rue Buffon, 75005 Paris, France,