1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, 100044, China
2. Department of Biological Sciences, University of Calgary, Calgary, AB T2N 1N4, Alberta, Canada,
3. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing, 100044, China,
4. Division of Paleontology, American Museum of Natural History, New York, 10024, New York, U.S.A.,
5. Earth and Environmental Sciences, Graduate Center, City University of New York, New York, 10017, New York, U.S.A.