1. Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, U.K., ,
2. Key Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology & Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China,
3. CAS Center of Excellence in Life and Paleoenvironment, Beijing, 100044, China
4. Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, U.K.,