1. Evolutionary Studies Institute and School of Geosciences, University of the Witwatersrand, Johannesburg, South Africa,
2. North Carolina Museum of Natural Sciences, 11 W. Jones Street, Raleigh, North Carolina 27601, U.S.A
3. Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg, South Africa
4. School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham B15 2TT, U.K.
5. Unidad Ejecutora Lillo, CONICET-Fundación Miguel Lillo, Miguel Lillo 251, San Miguel de Tucumán 4000, Tucumán, Argentina