1. Centre for Integrative Anatomy, Department of Cell and Developmental Biology, UCL, Anatomy Building, Gower Street, London WC1E 6BT, U.K.,
2. School of Earth Sciences, University of Bristol, Life Sciences Building, 24 Tyndall Avenue, Bristol BS8 1TQ, U.K.,
3. University Museum of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, U.K.