1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China,
2. Division of Earth Environmental System Science, Pukyong National University, Busan 48513, South Korea,
3. School of Earth and Environmental Sciences, Seoul National University, Seoul 08826, South Korea
4. Centre for Vertebrate Evolutionary Biology, Yunnan University, Kunming 650091, China