1. Instituto de Biodiversidad Neotropical, CONICET, Universidad Nacional de Tucumán, Crisóstomo Alvarez 722, San Miguel de Tucumán, Tucumán, Argentina, ;
2. Unidad Ejecutora Lillo, CONICET, Fundación Miguel Lillo, Miguel Lillo 251, San Miguel de Tucumán, Tucumán, Argentina;
3. Evolutionary Studies Institute, University of the Witwatersrand, 1 Jan Smuts Avenue, Braamfontein 2000, Johannesburg, South Africa, ;
4. Institute of Evolutionary Biology and Morphology, Center for Biomedical Education and Research, Faculty of Health, School of Medicine, University Witten/Herdecke, Alfred-Herrhausen-Strasze 50, 58448 Witten, Germany,
5. Albany Museum and Geology Department, Rhodes University, 40 Somerset Street, Grahamstown 6139, South Africa,