1. School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, B15 2TT, U.K.,
2. Department of Geosciences, Virginia Tech, Blacksburg, Virginia 24061, U.S.A.,
3. Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, U.K.,
4. Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, U.K.,