1. Korean Lichen Research Institute, Sunchon National University, Suncheon, South Korea;
2. Department of Plant Medicine, Sunchon National University, Suncheon, South Korea;
3. Korea National Arboretum, Gwangneungsumogwon-ro, Pocheon-si, Gyeonggi-do, South Korea;
4. Department of Natural Resources, Faculty of Applied Sciences, Sabaragamuwa University of Sri Lanka, Belihuloya, Sri Lanka