1. Department of Forest Science, Chungbuk National University, Cheongju, Republic of Korea;
2. Department of Biology, Chungbuk National University, Cheongju, Republic of Korea;
3. Division of Applied Life Science and Research Institute of Life Science, Gyeongsang National University, Jinju, Republic of Korea;
4. Department of Industrial Plant Science and Technology, Chungbuk National University, Cheongju, Republic of Korea