1. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, CAS, Kunming, China;
2. School of Life Science, Yunnan University, Kunming, China;
3. Korean Lichen Research Institute (KoLRI), Sunchon National University, Suncheon, Korea;
4. Snow and Landscape Research (WSL), Federal Institute for Forest, Zurich, Switzerland;
5. University of Bern, Bern, Switzerland