1. School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne, United Kingdom
2. School of Life Sciences, University of Essex, Colchester, United Kingdom
3. Technology & Innovation Delivery, Marine, Protective and Yacht, AkzoNobel/International Paint Ltd, Felling, Gateshead, United Kingdom
4. Physical Sciences Group, Platform Systems Division, Defence Science and Technology Laboratory (DSTL), Porton Down, Salisbury, United Kingdom