1. a Institute of Ecology and Evolutionary Biology , National Taiwan University , No.1, Sec. 4, Roosevelt Road, Taipei 106, Taipei , Taiwan
2. b Marine Biology Section, Department of Biology , University of Copenhagen , Universitetsparken 4, DK-2100, Copenhagen , Denmark
3. c Biodiversity Research Center , Academia Sinica , 128 section 2, Academia Road, Taipei 115, Taipei , Taiwan