1. Université de Toulon, MAPIEM-EA 4323, Toulon, France;
2. Coastal and Freshwater Group, Cawthron Institute, Private Bag 2, Nelson 7042, New Zealand;
3. Institute of Marine Science, University of Auckland, Private Bag 349, Warkworth 0941, New Zealand;
4. Université de Toulon, PROTEE-EA 3819, Toulon, France;
5. Université de Bretagne Sud, LBCM-EA 3883, IUEM, Lorient, France