1. a Departement de Biologie , Universite Cadi Ayyad , Marrakech, Maroc
2. b Laboratoire Sols et Environnement , ENSAIA , Vandoeuvre, France
3. c Department of Environmental Engineering Sciences , University of Florida , Gainesville, FL
4. d INRA Unite Agronomie , Centre de Recherches de Bordeaux , Villenave, France