1. a Scripps Institution of Oceanography, University of California , San Diego , California , USA
2. b Laboratory of Molecular Systematics, The Natural History Museum of Denmark , Copenhagen , Denmark
3. c Département Systématique et Evolution, Muséum National d'Histoire Naturelle , UMR 7205, Paris , France
4. d PROTEE (EBMA), Université du Sud – Toulon-Var , La Garde , France