1. Institut de Systématique, Évolution, Biodiversité (ISYEB), UMR 7205 CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, Paris, France;
2. University of Tübingen, Department of Evolution and Ecology, Auf der Morgenstelle 28E, Tübingen, Germany;
3. Findorffstr. 11, Ritterhude, Germany;
4. Department of Biology, Ithaca College, Ithaca, New York, USA