1. Wildlife Ecology and Management, University of Freiburg, Tennenbacher Straße 4, 79106 Freiburg, Germany
2. Institute of Wildlife Biology and Game Management, University of Natural Resources and Life Sciences Vienna, Gregor-Mendel Strasse 33, A-1180 Vienna, Austria.
3. Research Unit of Behavioural Ecology, Ethology and Wildlife Management, Department of Life Sciences, University of Siena, Via P.A. Mattioli 4, 53100 Siena, Italy.
4. Maremma Natural History Musem, Strada Corsini 5, 58100 Grosseto, Italy