1. School of Agriculture and Food Sciences, The University of Queensland, Gatton, Queensland 4343, Australia
2. Department of National Parks, Sport and Racing, Queensland Parks and Wildlife Service, Fraser Island, Queensland 4581, Australia
3. Fraser Coast Veterinary Services, Maryborough, Queensland 4650, Australia
4. Institute for Agriculture and the Environment, The University of Southern Queensland, Toowoomba, Queensland 4350, Australia