1. Graduate School of Environmental Earth Science, Hokkaido University, Kita 10, Nishi 5, Kita-ku, Sapporo 060-0810, Japan
2. Department of Evolutionary Studies of Biosystems, School of Advanced Sciences, Sokendai, The Graduate University for Advanced Studies, Hayama, Miura, Kanagawa 240-0193, Japan
3. Department of Biology and Geosciences, Graduate School of Science, Osaka City University, 3-3-138 Sugimoto, Sumiyoshi, Osaka 558-8585, Japan
4. Department of Biology, School of Biological Sciences, Tokai University, Minamisawa 5-1-1-1, Minami-ku Hokkaido, Sapporo 005-8601, Japan
5. Yamashina Institute for Ornithology, Konoyama 115, Abiko, Chiba 270-1145, Japan
6. Department of Natural History Science, Graduate School of Science, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060-0810, Japan