1. School of the Earth Sciences and Resources, China University of Geosciences, Beijing, China
2. Graduate School of Life and Environmental Sciences, University of Tsukuba, Ibaraki, Japan
3. School of Museum Studies, University of Leicester, University Road, Leicester, UK
4. Paleoscience Research Centre, University of New England, Armidale, Australia
5. Institute of Vertebrate Paleontology and Collage of Life Science and Technology, Gansu Agricultural University, Lanzhou, China