1. Institute of Geology and Paleontology, Linyi University, Linyi, China
2. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing, China
3. Graduate School, University of Chinese Academy of Sciences, Beijing, China
4. Department of Earth Sciences, The University of Hong Kong, Hong, Hong Kong