1. College of Resources and Civil Engineering, Northeastern University, Shenyang, China
2. Key Laboratory for Evolution of Past Life and Change of Environment, Liaoning Province, College of Palaeontology, Shenyang Normal University, Palaeontological Museum of Liaoning, Shenyang, China
3. Key Laboratory for Evolution of Past Life and Environment in Northeast Asia, Ministry of Education, Research Center of Palaeontology and Stratigraphy, Jilin University, Changchun, China
4. Directorate Earth and History of Life, Royal Belgian Institute of Natural Sciences, Brussels, Belgium