1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China;
2. College of Earth Science and Engineering, Shandong University of Science and Technology, Qingdao, China;
3. CAS Center for Excellence in Life and Paleoenvironment, Beijing, China;
4. University of Chinese Academy of Sciences, Beijing, China;
5. Research Center of Natural History and Culture, Qujing Normal University, Qujing, Yunnan, China