1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China
2. School of the Earth Sciences and Resources, China University of Geosciences (Beijing), Beijing, China
3. CAS Center for Excellence in Life and Paleoenvironment, Beijing, China
4. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing, China