1. State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Wuhan, China
2. Department of Animal Science, Hartpury College, Hartpury, UK
3. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing, China