1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China
2. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing, China
3. CAS Center for Excellence in Life and Paleoenvironment, Beijing, China
4. Division of Paleontology, American Museum of Natural History, New York, NY, USA