1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China
2. CAS Center for Excellence in Life and Paleoenvironment, Beijing, China
3. University of Chinese Academy of Sciences, Beijing, China
4. Division of Paleontology, American Museum of Natural History, New York, NY, USA
5. Richard Gilder Graduate School, American Museum of Natural History, New York, NY, USA