1. School of Earth Science and Engineering, Shandong University of Science and Technology, Qingdao, Shandong, China
2. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China
3. Deptartment of Earth Sciences, The Natural History Museum, London, UK
4. University of Chinese Academy of Sciences, Beijing, China