1. Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology Yunnan University, Kunming, China
2. MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Yunnan University, Kunming, China
3. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology Chinese Academy of Sciences, Nanjing, Jiangsu, China
4. University of Chinese Academy of Sciences, Beijing, China
5. Department of Earth Sciences, The Natural History Museum, London, UK