1. Key Laboratory of Forest Protection of National Forestry and Grassland Administration, Institute of Forest Ecology, Environment and Nature Conservation, Chinese Academy of Forestry, Beijing, China
2. College of Life Sciences, Capital Normal University, Beijing, China
3. Paleontological Institute, Russian Academy of Sciences, Russia
4. Department of Palaeontology, Natural History Museum, London, UK
5. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, Dc, USA